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Ecosystems and the Physical Environment
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PHYSICAL ENVIRONMENT
THE PHYSICAL ENVIRONMENT
We have seen how
living things depend on the physical environment to supply essential
materials for biogeochemical cycles. Physical factors such as climate
and soil also affect living things. (See Focus On: The Gaia Hypothesis
for an intriguing view of living organisms and their abiotic
environment.) Climate comprises the average weather conditions that
occur in a place over a period of years. Factors that determine an
area's climate include temperature, precipitation, wind, humidity, fog,
and cloud cover. Day-to-day variations, day-tonight variations, and
seasonal variations in these factors are also important aspects of
climate which we discuss in the remainder of this chapter, (Chapter 14
discusses soil, the surface layer of Earth that supports plants and is
home to countless numbers of bacteria, fungi, protists, and animals.)
The Sun Warms the Earth
The sun makes all life on Earth possible.
It warms the planet to habitable temperatures. Without the sun's energy,
the temperature on planet Earth would approach absolute zero (-273°C)
and all water would be frozen, even in the oceans. The hydrologic cycle,
carbon cycle, and other biogeo-chcmical cycles are powered by the sun,
and it is the primary determinant of Earth's climate. The sun's energy
is captured by photosynthetic organisms, which use it to make the food
molecules required by almost all forms of life. Most of our fuels wood,
oil, coal, and natural gas, for example— represent solar energy captured
by photosynthetic organisms. Without the sun, life on planet Earth would
cease.
The sun's energy is the product of a
massive nuclear fusion reaction (see Chapter 11) and is emitted into
space in the form of electromagnetic radiation—especially visible light
and infrared and ultraviolet radiation (which are not visible to the
human eye) An infinitesimal portion of this energy—one-billionth of the
sun's total production strikes the Earth's atmosphere, and of this
tiny-trickle of energy a minute part operates the eco-sphere.
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In the
daytime, 30 percent of the solar radiation that falls upon Earth is
immediately reflected away by clouds and surfaces, especially
snow, ice, and oceans. The remaining 70 percent is absorbed by the
Earth, where it runs the water cycle, drives winds and ocean
currents, powers photosynthesis, and warms the planet. Ultimately,
however, all of this energy is lost by the continual radiation of
long-wave infrared (heat) energy into space.
The
foregoing values are averages for the entire Earth and vary
substantially at different places
because of local conditions. For example,
high clouds increase energy reflection, whereas low clouds
increase energy absorption. |
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Solar Energy
at the Equator and the Poles
The
most significant local variation in
Earth's temperature is produced because the sun's energy doesn't reach
all places on Earth uniformly. A combination of the Earth's roughly
spherical shape and the tilt of its axis produce a great deal of
variation in the exposure of the Earth's surface to the energy
delivered by sunlight.
The principal
effect of the tilt is on the angles at which the sun's rays strike
different areas of the Earth at any one time. On the average, the sun's
rays hit the Earth vertically near the equator, making the energy more
concentrated and producing higher temperatures. Near the poles the sun's
rays hit more obliquely, and as a result their energy is spread over a
larger surface area. Also, rays of light entering the atmosphere
obliquely near the poles must pass through a deeper envelope of air than
those entering near the equator. This causes more of the sun's energy to
be scattered and reflected back to space, which in turn further lowers
temperatures near the poles. Thus, because the solar energy that reaches
Polar Regions is less concentrated, temperatures are lower.
Seasonal
Variations in Solar Energy
Seasons are
determined by two main factors: the inclination of the Earth's axis (the
more important factor) and the distance of the Earth from the sun, which
varies during the year. Since the Earth's inclination on its axis is
always the same (23.5°), during half of the year (March 21 to September
22) the Northern Hemisphere tilts toward the sun, and during
Hit-other half (September 22 to March 21) it tilts away from the sun.
(The orientation of the Southern Hemisphere is just the opposite at
these times.)
Atmospheric
Circulation
In large
measure, differences in temperature caused by variations in the amount
of solar energy reaching the Earth at different locations drive the
circulation of the atmosphere. The very warm surface of the Earth near
the equator heats the air that is in contact with it, causing this air
to expand and rise. As the warm air rises it cools, and then it sinks
again. Much of it recirculates almost immediately to the same areas it
has left, but the remainder of the heated air flows toward the poles,
where eventually it is chilled. Similar upward movements of warm air and
its subsequent flow toward the poles occur at higher latitudes (farther
from the equator) as well. As air cools by contact with the polar ground
and ocean, it sinks and flows toward the equator, generally beneath the
sheets of warm air that simultaneously flow toward the poles. The
constant motion of air transfers heat from the equator toward the poles,
and as the air returns, n cools the land over which it passes. This
continuous turnover does not equalize temperatures over the surface of
the Earth, but it does moderate them.
Surface Winds
In addition to
global circulation patterns, the Earth's atmosphere exhibits complex
horizontal movements that are commonly referred to as winds. The
nature of wind, with its turbulent gusts, eddies, and lulls, is
difficult to understand or predict. It results in part from differences
in atmospheric pressure and from the rotation of the Earth.
The gases that
constitute the atmosphere have weight and exert a pressure that is, at
sea level, about 1,013 millibars (14-7 pounds per square inch). Air
pressure is variable, however, changing with altitude, temperature, and
humidity. Winds tend to blow from areas of high atmospheric pressure to
areas of low pressure, and the greater the difference between the high-
and low-pressure areas, the stronger the wind.
The Earth's
rotation also influences the direction of wind. The Earth's rotation
from west to cast causes moving air to he deflected from its path and
swerve to the right in the Northern Hemisphere and to the left in the
Southern Hemisphere. This tendency is known as the Coriolis effect.
The Coriolis
effect can be visualized by imagining that you and a friend are
standing about 10 feet apart on a merry-go-round that is turning
clockwise. Suppose you throw a ball directly at your friend. By the
time the ball reaches the place where your friend was, he or she is no
longer in that spot. The ball will have swerved far to the left of your
friend. This is how the Coriolis effect works in the Southern
Hemisphere.
To visualize
how the Coriolis effect works in the Northern Hemisphere, imagine you
and your friend arc standing on the same merry-go-round, only this time
it is moving counterclockwise. Now when you throw the ball, it will
swerve far to the right of your friend.
The Earth's
atmosphere has three prevailing winds—major surface winds that blow more
or less continually. Prevailing winds that blow from the northeast near
the North Pole or from the southeast near the South Pole ate called
polar easterlies. Winds that blow in the mid-latitudes from the
southwest {in the Northern Hemisphere or from the northwest (in the
Southern Hemisphere) are called westerlies. Tropical winds that blows
from the northeast (Northern Hemisphere) or the southeast (Southern
Hemisphere) are called trade winds.
Patterns of
Circulation in the Oceans
The persistent
prevailing winds blowing over the ocean produce mass movements of
surface ocean water known as currents. The prevailing winds generate
circular ocean currents called gyres. For example, in the North
Atlantic, the tropical trade winds tend to blow toward the west, whereas
the westerlies in the mid-latitudes blow toward the east. This helps
establish a clockwise gyre in the North Atlantic. Thus, surface ocean
currents and winds tend to move in the same direction, although there
are many variations on this general rule. Other factors that contribute
to ocean currents include the Coriolis effect, the varying density of
water, and the positions of land masses.
The paths
traveled by surface ocean currents are partly caused by the Coriolis
effect . The Earth's rotation from west to east causes surface ocean
currents to swerve to the right in the Northern Hemisphere, creating a
circular, clockwise pattern of water currents. In the Southern
Hemisphere, ocean currents swerve to the left, thereby moving in a
circular, counterclockwise pattern.
The varying
density (mass per unit volume) of seawater affects deep ocean currents.
Water that is colder is denser than warmer water.' Thus, colder ocean
water sinks and flows under warmer water, creating currents far below
the surface. Deep ocean currents often travel in different directions
and at different speeds than do surface currents, in part because the
Coriolis effect is more pronounced at greater depths.
The positions of
land masses also affect oceanic circulation. As you can observe that the
oceans are not distributed uniformly over the globe: there is clearly
more water in the Southern Hemisphere than in the Northern Hemisphere.
Therefore, the circumpolar (around the pole) flow of water in the
Southern Hemisphere is almost unimpeded by land masses.
What Causes
Climate?
Average
temperature, temperature extremes, precipitation, the seasonal
distribution of precipitation, day length, and season length are the
most important dimensions of climate that affect living organisms.
Variations in these climatic factors produce the Earth's major
ecosystems, including tundra, desert, rain forest, and grassland.
Latitude
(distance north or south of the equator) and the inclination of the
Earth on its axis determine day length, season length, and, to a large
degree, temperature.
Differences in
precipitation depend upon several factors. The heavy rainfall of some
areas of the tropics results mainly from the equatorial upwelling of
moisture-laden air. High surface-water temperatures cause the
evaporation of vast quantities of water from tropical oceans, and
prevailing winds blow the resulting moist air over land masses. Heating
of the air by land surface that has been warmed by the sun causes moist
air to rise. As it rises, the air cools, and moisture condenses from
water vapor to a liquid, then falls as precipitation. The air eventually
returns to Earth on both sides of the equator between the Tropics of
Cancer and Capricorn (latitudes 23.5° north and 23.5° south). By then
most of its moisture has precipitated, and the dry air returns to the
equator. This air makes little biological difference over the ocean,
but its lack of moisture produces some of the great tropical
deserts, such as the Sahara Desert.
Air is also
dried by long journeys over land masses. Near the windward (the side
from which the wind blows) coasts of continents, rainfall may be heavy.
However, in the temperate zones—the areas between the tropics and the
polar zones— continental interiors are usually dry, because they are far
from oceans that replenish water in the air passing over them.
Moisture is also
removed from air by mountains when they cause humid air masses to rise
and thus
The El
Nino-Southern Oscillation Event and the World's Climate
Seasonal weather
forecasting requires an understanding of not only how the atmosphere
operates but also how the oceans interact with the atmosphere. Consider
the El Nino-Southern Oscillation event, a periodic warming of surface
waters of the tropical East Pacific that alters both oceanic and
atmospheric circulation patterns and results in unusual weather in
remote areas of the Earth. Every three to seven years, a warm mass of
water that is normally restricted to the western Pacific (near
Australia) expands eastward,
Increasing
surface temperatures in the east Pacific to 3 to 4 degrees over normal.
Ocean currents, which normally flow westward in this area, slow down,
stop altogether, or even
Reserve and go
eastward. The phenomenon is called El Nino (Spanish for "the child")
because the warming usually reaches the fishing grounds off Peru just
before Christmas.
The El
Nino-Southern Oscillation has a devastating effect on the fisheries off
South
America. The
higher temperatures and accompanying changes in ocean circulation
patterns prevent nutrient-laden deeper waters from upwelling {coming to
the surface). This severely decreases the populations of anchovies and
other marine organisms. During the 1972 El Nino, for example, the
anchovy population decreased by 90 percent.
The El
Nino-Southern Oscillation also alters air currents, directing unusual
weather to areas far from the tropical pacific. The 1991 El Nino, for
example, resulted in a much warmer-than-usual winter across much of
Alaska, western Canada, and the northern United States. El Nino was
responsible for the torrential rains that hit Texas and southern
California during the 1991-1992 winter season. In addition, the effects
of El Nino have been linked to droughts in Africa, Australia, and
Hawaii.
release their
water as precipitation. If prevailing winds blow onto a mountain range,
precipitation occurs primarily on the windward slopes of the mountains.
This situation exists on the west coast of
North America where precipitation falls
on the western slopes of the mountains. Downwind (in this case, east of
the mountain range), a low-precipitation rain shadow develops,
often creating a desert. Thus, some of the regional differences in
worldwide precipitation result from the drying of air as it is returned
to more equatorial areas; some result from long travel over
continents; and some
result from cooling produced by mountainous regions.
Variations in Overall Climatic
Conditions
Differences in elevation, in the
steepness of slopes and the directions they face, or in exposure to
prevailing winds may produce local variations in climate known as
microclimates, which can be quite different from the overall
climate surrounding them. The microclimate of an organism's habitat is
the climate it actually experiences and with which it must cope.
Sometimes it is possible for a population
of organisms to substantially modify its own microclimate, making it
more favorable. For example, trees modify the local climate within a
forest so that the temperature is usually lower, and the relative
humidity higher, than outside the forest. Beneath the litter of the
forest floor, temperature and humidity differ still more; the bottom of
the litter is cooler and moister than the surrounding forest. As
another example, desert-dwelling organisms burrow in the sand to evade
surface climatic conditions that would kill them in minutes. The cooler
daytime microclimate in their burrows permits them to survive until
night, when the surface cools off and they can leave their retreats to
forage or hunt.
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